Mario Bros. features two plumbers, Mario and Luigi, having to investigate the sewers of New York after strange creatures have been appearing down there. The objective of the game is to defeat all of the enemies in each phase. Both sides of every phase feature a mechanism that allows the player to go off-screen to the left and appear on the right and vice versa.
Mario Bros. features a scoring system in which points can be earned in several ways, such as by collecting coins or defeating enemies. The player gains points by defeating multiple enemies consecutively and can participate in a bonus round to gain more. Enemies are defeated by running up to them and kicking them when they are flipped on their back. Player cause enemies to flip by hitting them from below the platform they are on or by hitting the POW block to overturn all enemies on the ground. If the player allows the enemy to get up, the enemy becomes angry and increases in speed. Each phase has a certain number of enemies, ended with a more powerful enemy. Enemies come in variants—for example, Sidesteppers, a type of crab, are usually red, but turn blue and become faster when they recover from being flipped or if they are the final enemy. Each enemy has their own mechanics—the Shellcreeper, a type of turtle closely related to Koopa Troopas, can be made vulnerable by hitting them from below once, the Fighter Fly, a type of fly, jumps into the air every once in a while and must be hit from below when they are not in the air, and the Sidesteppers must be hit twice from below in that fashion to become vulnerable. Another enemy is called a Freezie, which has the ability to melt down and freeze the platform it is on, making it more difficult for the player to control the characters.
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